Grouped barplot in R with error bars

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Starting from your definition of df, you can do this in a few lines:

library(ggplot2)

cols = c(2,3,4,5)
df1  = transform(df, mean=rowMeans(df[cols]), sd=apply(df[cols],1, sd))

# df1 looks like this
#   Gene count1 count2 count3 count4 Species  mean        sd
#1 Gene1     12      4     36     12       A 16.00 13.856406
#2 Gene2     14      7     22     24       A 16.75  7.804913
#3 Gene1     16      9     54     35       B 28.50 20.240224
#4 Gene2     34     23     12     23       B 23.00  8.981462

ggplot(df1, aes(x=as.factor(Gene), y=mean, fill=Species)) +
  geom_bar(position=position_dodge(), stat="identity", colour="black") +
  geom_errorbar(aes(ymin=mean-sd, ymax=mean+sd), width=.2,position=position_dodge(.9))

enter image description here

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